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To measure iron concentration in whole-cell lysates, pelleted cells were lysed in optimized 2 RIPA lysis buffer (20 mM Tris-HCl, pH 7.4, 300 mM NaCl, 0.2% SDS, 0.1% sodium deoxycholate, 2% Triton X-100, 0.1 M HCl) for 20 min on ice, followed by centrifugation at 10,000 g for 10 min at 4 C

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